If your rim is hub centric then when you tighten the lugs the rim as centered. Yes, for the most part the load is carried by the lungs being tight, but that has jack-all to do with being centered on the axle. ...
On a 5 lug wheel, once the lugs are tight and the coned nuts are fully seated and centered in their respective holes, what more can the hub do beyond provide a 6th interference point for the five lugs to
potentially exert a force against? The "center" will always be the average of all the forces working together. Unless the hub is an interference fit into the wheel (be thankful it isn't), then the lugs will always have the ability to shift the wheel within the tolerance of the hub and wheel.
This is a very similar discussion as axle side track bar bolts and tapered tie rod end bolts. These parts are held in place by the clamping force of the properly torqued bolts. In this case, the cone shape of the lug nuts locate the final position of the wheel before hub does.
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